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Studies in the Psychology of Sex, Volume 1 The Evolution...

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Melinaud ("Pourquoi Rougit-on?" Revue des Deux Mondes, 1 Octobre, 1893) points out that blushing is always associated with fear, and indicates, in the various conditions under which it may arise,—modesty, timidity, confusion,—that we have something to conceal which we fear may be discovered. "All the evidence," Partridge states, "seems to point to the conclusion that the mental state underlying blushing belongs to the fear family. The presence of the feeling of dread, the palpitation of the heart, the impulse to escape, to hide, the shock, all confirms this view."
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G. Stanley Hall, "A Study of Fears," American Journal Psychology, 1897.
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Men are also very sensitive to any such inquisitiveness on the part of the opposite sex. To this cause, perhaps, and possibly, also, to the fear of causing disgust, may be ascribed the objection of men to undress before women artists and women doctors. I am told there is often difficulty in getting men to pose nude to women artists. Sir Jonathan Hutchinson was compelled, some years ago, to exclude lady members of the medical profession from the instructive demonstrations at his museum, "on account of the unwillingness of male patients to undress before them." A similar unwillingness is not found among women patients, but it must be remembered that, while women are accustomed to men as doctors, men (in England) are not yet accustomed to women as doctors.
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"I am acquainted with the case of a shy man," writes Dr. Harry Campbell, in his interesting study of "Morbid Shyness" (British Medical Journal, September 26, 1896), "who will make himself quite at home in the house of a blind person, and help himself to wine with the utmost confidence, whereas if a member of the family, who can see, comes into the room, all his old shyness returns, and he wishes himself far away."
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Stanley Hall ("Showing Off and Bashfulness," Pedagogical Seminary, June, 1903), quotes Dr. Anagnos, of the Perkins Institute for the Blind, to this effect.
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Thus, Sonnini, in the eighteenth century, noted that the country women in Egypt only wore a single garment, open from the armpits to the knees on each side, so that it revealed the body at every movement; "but this troubles the women little, provided the face is not exposed." (Voyage dans la Haute et Basse Egypte, 1779, vol. i, p, 289.) When Casanova was at Constantinople, the Comte de Bonneval, a convert to Islam, assured him that he was mistaken in trying to see a woman's face when he might easily obtain greater favors from her. "The most reserved of Turkish women," the Comte assured him, "only carries her modesty in her face, and as soon as her veil is on she is sure that she will never blush at anything." (Mémoires, vol. i, p. 429.)
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It is worth noting that this impulse is rooted in the natural instinctive acts and ideas of childhood. Stanley Hall, dealing with the "Early Sense of Self," in the report already mentioned, refers to the eyes as perhaps even more than the hands, feet, and mouth, "the centres of that kind of self-consciousness which is always mindful of how the self appears to others," and proceeds to mention "the very common impression of young children that if the eyes are covered or closed they cannot be seen. Some think the entire body thus vanishes from sight of others; some, that the head also ceases to be visible; and a still higher form of this curious psychosis is that, when they are closed, the soul cannot be seen." (American Journal of Psychology, vol. ix, No. 3, 1898.) The instinctive and unreasoned character of this act is further shown by its occurrence in idiots. Näcke mentions that he once had occasion to examine the abdomen of an idiot, who, thereupon, attempted to draw down his shirt with the left hand, while with the right he covered his eyes.
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Cf. Stanley Hall and T. Smith, "Showing Off and Bashfulness," American Journal of Psychology, June, 1903.

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Studies in the Psychology of Sex, Volume 1 The Evolution of Modesty; The Phenomena of Sexual Periodicity; Auto-Erotism, Project Gutenberg eBook #13610

III. IV.